Mediterranic.com - Deep time & dedication. Moroccan Fossils, Scientifically Curated since 2004. Free papers. 
👉 Scroll to Shop

Museum-quality Moroccan trilobites with scientific papers. Browse specimens, request fossils, and bid on auctions. 4.98/5 ⭐ rated seller.

Moroccan Trilobites — Fossils, Papers & Shop | Linktree

eBay listings

NEW ARRIVALS + Specimen Request & Shipping Quote

See Why Collectors Trust Us (4.98/5 ⭐)

Shop: All Listings

DEVONIAN TRILOBITES

CAMBRIAN TRILOBITES

Catawiki auctions

📚 Free Papers (Download PDFs)

Website (About + Gallery)

Gerd Geyer (2020).

Abstract
A characteristic group of redlichioid trilobites distributed in the late part of Cambrian Stage 3 and the early part of Cambrian Stage 4 in West Gondwana is revised and interpreted to constitute a monophyletic family Despujolsiidae. More than 25 species were previously assigned to the genera Perrector, Resserops, Eops, Marsaisia, Despojulsia, Pareops, Clariondia, Realaspis and Dolerolichia and are critically evaluated and revised herein, with Resserops regarded as an objective synonym of Perrector, and Pareops as being tentatively a synonym of Marsaisia. Species assigned to Resserops and Pseudoresserops reported from Jordan, the Siberian Platform and Mongolia are shown to belong to other clades. The species of Pararedlichia and the concept of the subfamily Pararedlichiinae and the family Abadiellidae, as well as relationships to and within the family Dolerolenidae are critically evaluated. The study includes data on relics of the digestive tract seen in specimens of Perrector falloti and P. brevilimbatus from strata of the Daguinaspis Zone of the western Anti-Atlas. They belong to the hitherto neglected Souss Fossil Lagerstaette and are the first published data on Cambrian trilobites with soft-part preservation from Africa. In addition, new taxa described from Morocco include Epicurella pustulata n. gen., n. sp., Marsaisia devoillei n. sp. and Marsaisia? lemdadensis n. sp. Moreover, Dolerolenus courtessolei Pillola, 1991 is regarded as representing a new genus, Boethiusia n. gen.

This is an open-source publication. Follow all the steps in your email (including signing our newsletter if you wich) and you will receive the download link.

Resserops clade: Despujolsiidae

Gerd Geyer (1996).

Abstract. Restudy of the type material and numerous additional specimens of Falllotaspis, Chouberte/la, and Daguinaspis from Morocco suggest several changes in the species-level taxonomy of these genera. Most of HUPE's (1953a) six species of Fallotaspis appear to represent valid species. However, Fallotaspis planospinosa is represented by a poorly preserved specimen, and the species is to be restricted to the holotype. Fallotaspis longispina HUPE, 1953, and F longa HUPE, 1953, are both junior synonyms of 0/ene/lus bondoni NELTNER & POCTEY, 1950, and the valid name is Fallotaspis bondoni (NELTNER & POCTEY, 1950). Choubertella lata HUPE, 1953, is a subjective junior synonym of C. spinosa HUPE, 1953, and C. crassioculata HUPE, 1953, is known only from the holotype, which probably represents a juvenile cephalon. Of HUPE's live species of Daguinaspis, D. latifrons HUPE, 1953, D. subabadiei HUPE, 1953, and D. subabadiei HuPE, 1953, are regarded as junior subjective synonyms of the type species, D. ambroggii HUPE & ABADIE, 1950. Daguinaspis (Epidaguinaspis) angusta HUPE, 1953, and D. (Eodaguinaspis) abadiei HUPE, 1953, are both only known from the holotypes and are dubious. Neither ofthe subgenera Eodaguinaspis and Epidaguinaspis appears tobe valid or required for a subdivision ofthe genus Daguinaspis, and they are rejected herein. Fallotaspis inexspectatus n. sp. is newly established.

The Siberian species Fallotaspis sibirica REPINA, 1965, and F explicata REPINA, 1965, represent the new genus Repinaella. Tagether with the genera Profa/iotaspis REPJNA, 1965, and Lenal/ina REPINA, 1990, earlier assigned to the Fallotaspidinae, it is assigned to the family Archaeaspididae.
The fallotaspidid trilobites Eojallotaspis, Fallotaspis, Choubertella, and Daguinaspis are closely related phylogenetically. As a con- sequence, their Stratigraphie occurrence gives some evidence for their order of descent. They are the index fossils of the lower !Our zones of the biostratigraphical zonation in southern Morocco and are indicative and characteristic of the Issendalen Stage. However, the genus Fallotaspis has a long stratigraphical range, which includes part of the Eojallotaspis Zone and does not define a limited time range useful for international correlations.

The olenelloid and redlichioid phylogeny is reevaluated. A number of morphologic aspects, as weil as the presence of a protaspid larva and the Stratigraphie occurrences, strongly suggest a systematic differentiation of fallotaspidid and olenellid trilobites. Fallotaspidid trilobites are regarded as the ancestors of the such redlichiids as Lemdadella. Olenellids and holmiids, in contrast, represent a different lineage that appears to have given rise to the Xystriduridae.

Important differences between Fallotaspis with Choubertel/a and Daguinaspis are due to the reduced width of the Cephalon in both Choubertella and Daguinaspis. As a result, extended genal spines are absent in Choubertella and Daguinaspis and indicate a modification of the ancestral body plan of Fallotaspis. The characters visible on the carapace present a mosaic of Features known from different olenelloid and redlichioid taxa and do not per se readily define the group. Details on the general morphology and its functional aspects, including aspects of the generat body plan, glabella, cephalic border and frontal area, eye orientation and size, genal ridges and »metaparian suture«, doublure, rostral plate, cephalic segmentation, thorax, pygidium, and cuticular ornament, are described in detail and considered in developing a phylogenetic systematics of these trilobites. The visual surface of the three genera is in a vertical position with its lower and upper margin parallel to the ventral surface of the cephalic border that lay in a horizontal position during life. investigation of intergenal ridges, posterior ocular lines, and associated fractures indicate that the "metaparian suture" of the fallotaspidids does not exist and this type of "suture" is mimiced by cracks, which commence at the posterior end of the visual surface. In Choubertella and Daguinaspis, the visual surface was considerably elevated above the cephalic border, and this Ieads to the assumption that Chouberte/la and Daguinaspis preferably lived on soft substrate as »mud creepers« . The relief o f the ocular lobe helped the animals keep the eyes beyond the sediment. In general, morphologic rcevaluation Ieads to an understanding of the differences between these species and genera and to elucidating evolutionary patterns among the Fallotaspididae, and to deterrnining important differences between them and the olenelloid and holmiid trilobites. Reconstruction ofexuviation supplies additional data.

The articulation of the thoracic segments of Fal/otaspis includes a combination of a flange joint with articulating processes at the axial furrows with fulcra and corresponding swellings ! that acted as limiting marginal connective devices. This articulation permitted an incomplete cylindrical enrollment. Additional discussions summarize facts on the autecology and synecology of Fallotaspis, Chouber- tel/a, and Daguinaspis. A protaspid larva of Fal/otaspis i described for the fir t time. lts shape suggests a benthic mode of life. A meraspid cephalon of Chouberte/la indicates both a well developed intergenal spine and a genal spine that is reduced to a rudimentary structure in adult individuals.

The sclerites of Fallotaspis, Chouberte/la, and Daguinaspis are usually affected by complex diagenetic as weil as taphonomic processes, and their composition differs largely from the original composition ofthe cuticles. In different lithofacies, the exoskeletal composition was specifically altered. The main effect on the mode of preservation results from different grain sizes ofthe siliciclastic Sediments. All of the specimens preserved in fine-grained sandstones and siltstones are composite-molds. ln the usual type of preservation in shale, the primary cuticular substances and fabrics were destroyed and repla ed by such mixed components as argillaceous, phosphatic, and ferritic minerals. Preservational features and the mode of deformation Iead to the assumption that the cuticle of Fallotnspis was slightly less calcified or that the cuticular calcite was more unstable by comparison with redlichiid or ellipsocephalid trilobites.

The circumocular suture in Eofallotaspis, Fallotaspis, Choubertella and Daguinaspis are frequently the locus ~fadditional frac.ture~, which tend to align with the major axes of the cephalon and resulting compactional vectors, a pheno~1enon th~t IS not pa~all~led m tn- l o b i t e s w i t h w e i l d e v e l o p e d f a c i a l s u t u r e . T h i s . i n t u r n , s h o w s t h e i m p o r t a n c e o f a w e i l ~eveloped .fac~al s u t u r e . m . I a r g e r t n l o b 1 t e s . Fr~c­ tures at the anterior end of thc eye and from the posterior end of the eyes to the postenor cephaltc nm are pnm1t1ve stages of a fac1al suture. Such fractures may hardly be distinguishable from »real« sutures. . . . . .

Most strata with specimens of Fallotaspis, Choubertelfa, and Daguinaspis contain isolated or artJculate~ remains of mdiVI~uals,
not show indication of any notable sorting. Hence, most assemblages appear to be undisturbed by any kmd of sedlmentologlcal or pred- ator/scavenger activity. Beds with abundant assemblages ofspecimens of Daguinaspis in the Amouslek and Taze.mmourt sections ar.e a remarkable exception, and sedimentological as weil as taphonomical facts suggest a pronounced post-martern s1ze- and shape- ort1ng due to lateral transport.
Quantitative statistical analyses were used to define the morphologic variation in the three genera Fallotaspis, Choubertella, and Daguinaspis. They include primarily regre sion analyses, as weil as multidimensional scaling. In addition, the wealth of data allow rein- vestigation ofearlier statistical tests on 0/enus from Sweden, which tumed outtobe apparently insignificant.

This is an open-source publication. Follow all the steps in your email (including signing up our newsletter if you wich) and you will receive the download link.

Fallotaspidid trilobites revisited (Geyer, 1996)

Geyer, G. 2019: The earliest known West Gondwanan trilobites from the Anti-Atlas of Morocco, with a revision of the Family Bigotinidae Hupe, 1953. Fossils and Strata, No. 64, pp. 55–153.

The previously poorly investigated trilobite fauna of the upper Igoudine and lowest Amouslek formations in the western Anti-Atlas, Morocco, critical for the understand-
ing of the earliest trilobites on a global scale, is studied in detail and its taxonomic diversity and biostratigraphical characteristics reviewed. The key section at Tiout, par-
ticularly the Tiout Member, yields surprisingly diverse faunal associations with trilo-
bites of the families Bigotinidae and Fallotaspididae, of which only Hupetina antiqua,
Eofallotaspis prima, E. tioutensis and Fallotaspis cf. F. tazemmourtensis have been pre-
viously identiﬁed. New genera and species from the Tiout section comprise Bigotina
kelleri n. sp., B. monningeri n. sp., Issendalenia grandispina n. gen., n. sp., Tioutella
ﬂoccofavosa n. gen., n. sp., Pseudobigotina antiatlasensis n. gen., n. sp., Eladiolinania
castor n. gen., n. sp., E. pollux n. gen., n. sp., Debrenella larvalis n. gen., n. sp. and Fal-
lotaspis antecedens n. sp. Bigotinops chouberti n. sp. is a new species from below the
known range of Fallotaspis tazemmourtensis in the Amouslek section. A revision of
other bigotinid genera and species from the Iberian sector of West Gondwana, Cado-
mian France, the Siberian Platform and the Altay-Sayan Foldbelt leads to a recombina-
tion of Bigotina angulata Suvorova, 1960; Eladiolinania? palaciosi Lin~an et al., 2008;
E.? gordaensis Lin~an et al., 2008; Suvorovaella priva (Suvorova, 1960) and Suvoro-
vaella? patria (Suvorova, 1960) and a suggested new family Minusinellidae. The previ-
ously established Eofallotaspis and Fallotaspis tazemmourtensis zones in the Moroccan
biostratigraphical scheme are revised to become the Hupetina antiqua, Eofallotaspis
tioutensis and Fallotaspis plana zones. □ Biostratigraphy, Lower Cambrian, phylogeny,
Sibiria, Trilobita, West Gondwana.

This is an open-source publication. Follow all the steps in your email and you will receive the download link.

Family Bigotinidae (Geyer, 2019)

Unexpectedly curved spines in a Cambrian trilobite: considerations on the spinosity in Kingaspidoides spinirecurvatus sp. nov. from the Anti‐Atlas, Morocco, and related Cambrian ellipsocephaloids

Gerd Geyer · Miguel Caldeira Pais · Thomas Wotte

Abstract

The new ellipsocephaloid trilobite species Kingaspidoides spinirecurvatus has a spectacular morphology because of a unique set of two long and anteriorly recurved spines on the occipital ring and the axial ring of thoracic segment 8. Together with the long genal spines this whimsical dorsally directed spine arrangement is thought to act as a non-standard protective device against predators. This is illustrated by the body posture during different stages of enrolment, contrasting with the more sophisticated spinosities seen in later trilobites, which are discussed in brief. Kingaspidoides spinirecurvatus from the lower–middle Cambrian boundary interval of the eastern Anti-Atlas in Morocco has been known for about two decades, with specimens handled as precious objects on the fossil market. Similar, but far less spectacular, spine arrangements on the thoracic axial rings are known from other ellipsocephaloid trilobites from the Anti-Atlas of Morocco and the Franconian Forest region of Germany. This suggests that an experimental phase of spine development took place within the Kingaspidoides clade during the early–middle Cambrian boundary interval.

Keywords Cambrian · Trilobita · Systematics · Biostratigraphy · West Gondwana · Morocco

This is an open-source publication. Follow all the steps in your email (including signing up to our Newsletter if you wish) and you will receive the download link.

Unexpectedly curved spines in a Cambrian trilobite: considerations on the spinosity in Kingaspidoides spinirecurvatus sp. nov.

Kingaspidoides spinirecurvatus

The Souss lagerstätte of the Anti‐Atlas, Morocco: discovery of the first Cambrian fossil lagerstätte from Africa (Geyer, G. & Landing, E. 2021)

Gerd Geyer & Ed Landing (2021).

Episodic low oxygenated conditions on the seafloor are likely responsible for the exceptional preservation of animal remains in the upper Amouslek Formation (lower Cambrian, Stage 3) on the northern slope of the western Anti‐Atlas, Morocco. This stratigraphic interval has yielded trilobite, brachiopod,
and hyolith fossils with preserved soft parts, including some of the oldest known trilobite guts. The “Souss fossil lagerstätte” (newly proposed designation) represents the first Cambrian fossil lagerstätte in Cambrian strata known from Africa and is one of the oldest trilobite‐bearing fossil lagerstätten

on Earth. Inter‐regional correlation of the Souss fossil lagerstätte in West Gondwana suggests its development during an interval of high eustatic levels recorded by dark shales that occur in informal upper Cambrian Series 2 in Siberia, South China, and East Gondwana.

This is an open-source publication. Follow all the steps in your email and you will receive the download link.

The Souss lagerstätte of the Anti‐Atlas

Gerd Geyer, Ed Landing, Miguel Caldeira Pais, Nasrrddine Youbi and Moulay Ahmed Boumehdi (2020)

The Tazemmourt section (Fig. 1) is composed of lower Cambrian strata with a total thickness of ca. 350 m. It includes rocks of the Igoudine and Amouslek formations. The section is fairly well studied (e.g., [1]–[6]), but is particularly famous for its superbly preserved trilobites (extinct marine arthropods with dorsal skeletons mineralized by calcium carbonate). Many trilobite species were described from the Tazemmourt section for the first time and have their type localities here ([1], [8]). In addition, the vertical sequence of fossils seen in the section is the basis for the biostratigraphical subdivision in the Atlas Ranges. This fossil-based zonation is part of the most important relative time subdivisions of the Cambrian Period in the West Gondwana paleocontinent and globally (e.g., [1], [6], [9]–[11]).

Recent work has revealed that several horizons in the Daguinaspis Zone (based on trilobite fossils) of the Amouslek Formation yield animal remains with well preserved, internal soft-parts. These occurrences were termed the Souss Lagerstätte, and this is the first Cambrian fossil Lagerstätte known from Africa ([12], [13]).

This is an open-source publication. Follow all the steps in your email and you will receive the download link.

The Tazemmourt Section (Geyer et al., 2020)

Tazemmourt Section (Geyer et al., 2020)

This document provides detailed paleontological research on a newly identified obolellid brachiopod species (Brevipelta chouberti - Geyer, 1994) discovered in the Lower Cambrian strata of Morocco, offering insights into ancient marine life and geological history.

ABSTRACT
A new genus and species of obolellid brachiopod, Brevipelta chouberti, is described from the Early Cambrian of Morocco.
It is a useful index fossil for the upper Choubertella Zone and the Daguinaspis Zone in Morocco. Additionally discussed are taphonomic
aspects that lead to the question whether the shell may have been weakly calcified compared to other obolellid brachiopods and the
calcareous part may have been composed of aragonite crystallites, or whether the preservation of Brevipelta is due to facile solubility
of a calcitic shell substance. Brevipelta chouberti represents not only the oldest described Early Cambrian brachiopod from Africa,
but one of the oldest obolellid brachiopods known worldwide.

Brevipelta chouberti (Gerd, G., 1994): A New Obolellid Brachiopod from the Lower Cambrian of Morocco

Brevipelta chouberti

The Souss Lagerstätte - MEDITERRANIC

ARRIVALS + Request Specimen Quote

Moroccan Trilobites — Fossils, Papers & Shop

Mediterranic

DEVONIAN

Comura bultynchi 5,6 cm Trilobite. Jbel Oufaten

Devonian trilobites

Longianda termieri -RARELY OFFERED 11.6 cm (!) – Issafen Cambrian Trilobite

TOP RARE Bigotinops dangeardi Fossil Trilobite Morocco Cambrian BIGOTINOIDS

Fallotaspis bondoni - SOUGHT-AFTER 1.4 cm – Souss Lagerstätte Trilobite - Fossilised animal

Fallotaspis tazemmourtensis - SOUGHT-AFTER 3.9 cm (!) – Souss Lagerstätte Trilobite - Fossilised animal

SOFT TISSUES 0.9 cm Bigotinops dangeardi – Souss Lagerstätte Trilobite - Fossilised animal

Cambrian Trilobites

Get Notified first of NEW ARRIVALS or Tell Me the species/type you want + your shipping country. I’ll reply with options and total price. Response time: ~ 24h.

Enjoy 30 days of Hulu for FREE

Claim 10 free meals* from HelloFresh

Get 60% off Sitewide

Sip your way to stronger health with 30% off at ARMRA

Save 10% at Omnilux LED

Save 10% on Ritual Multivitamins Subscriptions

You've scored 50% OFF Factor 🤤

Join now and get 50% off a Club membership

Get your first box FREE with Curology!

Claim your $20 bonus with Acorns

Sign up for Headspace Today and get a 14 day free trial!

Get AAA membership for half the price!

Get 30¢/gal cash back on gas at major stations

Get 15% off at Equip Foods

20% Off the at FabFitFun with code INSIDER20

Get 80% off your first order when signing up to be a VIP!

Get 15% off for cleaner skin at CLEARSTEM

Get your first box FREE with Agency!

50% off + Free Shipping at Purple Carrot!

Power Up with Daily Harvest's High-Protein Smoothies

Save $100 with Gobble

Save 10% on your First Maev Order!

15% off your favorite non-alcoholic spirits at The Zero Proof

10% off JLab’s award‑winning audio gear

Get help with or without insurance. Use code SPACE80 for $80 off.

Start speaking a new language in just 3 weeks

Mediterranic.com - Deep time & dedication. Moroccan Fossils, Scientifically Curated since 2004. Free papers. 👉 Scroll to Shop

📚 Free Papers (Download PDFs)

About this account

More from Linktree